Reproductive success measures how well birds produce
viable offspring. It can be calculated and interpreted in various
ways depending on the stage of the nesting cycle. Success rates
can offer insight about the most critical stage of the nesting cycle
and provide useful information for conservation efforts.
In Table 1, we summarize nest-box success rates for six of The
Birdhouse Network's most common species and six of our "Most
Wanted" birds. We used three different measures to help us
determine the stage when a species is most vulnerable to nest
failure. The nestling rate is the percentage of eggs that actually
hatched, and fledgling rate measures the percentage of eggs in
completed clutches that ultimately produce fledglings. For both
the nestling and fledgling rates, we considered only nests with
complete clutches and at least one nestling present. This enabled
us to measure parental ability in nests that made it to the nestling
stage. Overall nesting success is the proportion of all nest attempts
that produced at least one fledgling. This measure includes nests
with at least one egg recorded, even if the eggs never hatched.
| SPECIES |
# of Nest Attempts |
# of Complete
Clutches
|
Nestling Rate (%) |
Fledgling Rate (%) |
Overall Nesting Success
(%) |
Average Clutch Size |
Average # Nestings |
Average # Fledglings |
| Eastern Bluebird |
3,313
|
2,690
|
90
|
88
|
72
|
4.4
|
4.0
|
3.9
|
| Tree Swallow |
2,283
|
1,803
|
89
|
87
|
70
|
5.3
|
4.8
|
4.6
|
| House Wren |
737
|
555
|
90
|
90
|
65
|
5.6
|
5.2
|
5.1
|
| Mountain Bluebird |
597
|
518
|
88
|
88
|
78
|
5.0
|
4.5
|
4.4
|
| House Sparrow |
354
|
112
|
83
|
81
|
17
|
4.5
|
3.6
|
3.9
|
| Western
Bluebird |
311
|
275
|
87
|
83
|
77
|
5.1
|
4.5
|
4.3
|
| Purple Martin |
88
|
71
|
90
|
88
|
76
|
5.4
|
5.0
|
4.7
|
| American Kestrel |
44
|
34
|
96
|
93
|
80
|
4.6
|
4.4
|
4.2
|
| Mountain Chickadee |
36
|
32
|
91
|
88
|
81
|
6.3
|
5.8
|
5.7
|
| Bewick's Wren |
32
|
23
|
89
|
84
|
72
|
5.8
|
5.5
|
5.3
|
| Prothonotary Warbler |
30
|
22
|
73
|
73
|
67
|
4.9
|
3.6
|
3.7
|
| Great Creasted Flycatcher |
28
|
26
|
91
|
90
|
82
|
4.3
|
4.3
|
4.4
|
| Table 1.
Summary of 2000-01 nest data. The Birdhouse Network's
"Most Wanted" shaded in pink. # of Nest
Attempts = nests with at least one egg. # of
Complete Clutches = nests with at least one nestling.
Nestling Rate = % eggs hatched in nests with
at least one nestling. Fledgling Rate = % nests
with at least one nestling that produced at least one
fledgling. Overall Nesting Success = % nest attempts
producing at least one fledgling in nests that had at
least one egg. Average Clutch Size and Average
# Nestlings are based on # Complete Clutches. Average
# Fledglings is based on nests with at least one
fledgling, accounting for higher fledgling averages
than nestling averages. For other species, click
here. |
|
To discern what happens during the various stages of the nesting
cycle, choose a species from Table 1 and follow the rows from
nestling rate to overall nesting success. For example, the Tree
Swallow shows a nestling rate of 89 percent and a fledgling rate
of 87 percent. This suggests that once Tree Swallows make it to
the nestling stage, they have a high probability of fledging.
Now look at the overall nesting success rate, which includes nests
that never produced nestlings. The rate of 70 percent is lower
than the rates for more than half of the other birds listed and
suggests that Tree Swallows have a relatively high rate of failure
during egg laying and incubation. Note that Table 1 is sorted
from highest to lowest sample size and thus, highest to lowest
reliability.
The overall nesting success rate ranged from 65 percent (House
Wrens) to 82 percent (Great Crested Flycatchers), except for House
Sparrows, with an anomalously low 17 percent. Figure 1 indicates
why nests failed. Ninety-two percent of House Sparrow nest failures
were caused by human intervention, a practice both legal and encouraged
for nonnative species that usurp cavities from native birds. Among
the other species, the most common reason for nest failure was
predation, accounting for more than 40 percent of all failed nests
for Eastern Bluebirds, Tree Swallows, House Wrens, and Mountain
Chickadees. Nest abandonment was another major cause of failure,
accounting for at least 20 percent of failed nests for seven species.
|
|
| Figure 1. Causes of nest failures, 2000-01,
for nests with at least one egg but no fledglings reported.
Numbers in parentheses indicate number of failed nests. |
Providing a nest box is a great way to increase nesting sites
for cavity-nesting birds, but if it also increases the likelihood
of predation, guards should be erected. The Birdhouse Network
strongly encourages the use of predator guards, particularly in
areas where predators are abundant. In addition, regular, weekly
monitoring and recording of nest-box activity will help increase
our overall knowledge of nesting success and failure.
Suggested citation: Phillips, Tina
and Ronald Rohrbaugh. Measuring Their Success. Birdscope,
newsletter of the Cornell Lab of Ornithology, Winter 2002. <www.birds.cornell.edu>
For permission to reprint all or
part of this article, please contact Miyoko Chu, Editor, Cornell
Lab of Ornithology, 159 Sapsucker Woods Rd., Ithaca, New York.
Phone (607) 254-2451. Email mcc37@cornell.edu