Handbook of Bird Biology
Editors:
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Rick Bonney Rick has a B.S. in wildlife biology and a M.P.S. in natural resources extension, both from Cornell University, and he has been working at the Lab since 1981. A Cornell Senior Extension Associate, his research focuses on development and evaluation of inquiry-based science education projects. |
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Sandy Podulka The primary editor of the course, Sandy also developed the course exams. She received a B.S. in Wildlife Biology from Cornell University and an M.S. in Zoology (Animal Behavior) from the University of Maryland, where she studied the function of song repertoires in Song Sparrows. |
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Ron Rohrbaugh Second of the course editors and currently Acting Director of the Conservation Science Program, Ron guided the production of the course. He holds B.S. and M.S. degrees in Wildlife Biology from Penn State University. Ron's research focuses on the breeding biology of American Kestrels, grassland ecology, and using citizen science to study and monitor bird populations. |
Authors
Kenneth P. Able,
SUNY Albany
CHAPTER 5: Birds on the Move: Flight and Migration
Kenneth P. Able is Professor of Biology in the Department of
Biological Sciences at the State University of New York in Albany,
where he has been a member of the faculty since 1971. He received his
B.S. and M.S. degrees from the University of Louisville, and his Ph.D.
from the University of Georgia. Ken's research focuses on bird
migration, particularly the mechanisms of orientation and
navigation.
Ken has been passionately interested in birds since childhood. He has
birded extensively across North America and in Mexico, Costa Rica,
Peru, Spain, France, Germany, Italy, Switzerland, the United Kingdom,
Australia, New Zealand, and South Africa. His favorite destination is
Australia, because the avifauna is unique and the species
fantastic.
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John
Alcock, Arizona State University
CHAPTER 6: Understanding Bird Behavior
John Alcock is Regents' Professor of Biology at Arizona State
University, Tempe. John is currently researching the evolution of
insect mating systems, with a special emphasis on the diversity of male
mating tactics among bee species native to the Sonoran Desert. Earlier
research has focused on birds, however, and his textbook, Animal
Behavior, An Evolutionary Approach uses many bird examples to
illustrate all aspects of the modern study of behavior.
John began bird watching at age 5 (bird number one was the mallard),
and seeing a good bird still boosts his heart rate. In attempts to keep
his heart rate up, he has visited Costa Rica, Ecuador, Argentina,
Australia, and several countries in Europe.
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George A. Clark,
University of Connecticut
CHAPTER 3: Form and Function: The External Bird
George A. Clark, Jr. is Professor Emeritus of Ecology and
Evolutionary Biology at The University of Connecticut in Storrs. He
received his Ph.D. in Biology from Yale University, where he
specialized in ornithology. After spending two years at the University
of Washington in Seattle, he moved to The University of Connecticut
where he spent 32 years as a faculty member.
His 200 publications reflect his research interests in the structure,
behavior, distribution, and evolution of birds. He is past president of
the Association of Field Ornithologists, served as Co-Editor of the
book Perspectives in Ornithology/ Essays Presented for the
Centennial of the American Ornithologists' Union, and has led
educational field trips for groups to observe birds in North and South
America, Europe, and Africa.
A birder since his high school days in Pennsylvania, George now
resides in Vermont and enjoys seeking birds by walking or snowshoeing
in the northern New England hills.
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Marie Z.
Eckhardt, Cornell University
Birds and Humans: A Historical Perspective
Marie Eckhardt is a biologist, formerly employed with the Education
Department of the Cornell Lab of Ornithology. Her academic background
in vertebrate zoology and experience as a museum consultant and as
collections manager at the New York State Museum provided a strong
foundation for her varied contributions at the Lab. Marie has been
interested in animals and the natural world for as long as she can
remember—an interest she credits to early and regular museum
visits.
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Howard E. Evans, Cornell
University
CHAPTER 4: What's Inside: Anatomy and Physiology
Howard E. Evans received both his B.S. and Ph.D. in
comparative anatomy from Cornell University, where he became a faculty
member in the Veterinary College in 1950. There he taught gross anatomy
of the horse and cow for seven years, and anatomy of the dog, bird, and
fish for 36 years. He was secretary of the college for 12 years and
served as chairman of Anatomy from 1976 to 1986, when he retired. He
continues to teach a course on the literature and materials of natural
history, and to lecture in several other courses.
Howie's research concerned the anatomy of reptiles and birds, the
replacement of teeth in fishes, the plant-induced cyclopis in sheep,
fetal development of the dog, and anatomy of tropical fishes. His most
recent works include the third edition of Miller's Anatomy of the
Dog (1993), the fifth edition of Guide to the Dissection of
the Dog (2000 with Dr. de Lahunta), and the third edition of
Anatomy of the Budgerigar and Other Birds (1996). He is
co-editor of the Handbook of Avian Anatomy published by the
Nuttall Ornithological Club of Harvard University. He has served as
president of both the American and The World Association of Veterinary
Anatomists, and was an associate editor of the American Journal of
Anatomy and the Journal of Morphology.
A native of New York City, Howie and his wife Erica have led natural
history trips for the Cornell Adult University to the Virgin Islands;
Hawaii; Sapelo Island, Georgia; East and South Africa; Papua, New
Guinea; and Antarctica. He has lectured in China, Russia, Taiwan,
Hungary, Switzerland, Germany, England, Thailand, Brazil, Mexico, and
Japan.
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Alan
Feduccia, University of North Carolina
Evolution of Birds and Avian Flight
Alan Feduccia is S. K. Heninger Professor of Biology at the
University of North Carolina at Chapel Hill, where he has been for 30
years. He received his Ph.D. at the University of Michigan, where his
thesis focused on the evolution of woodhewers and ovenbirds.
Alan's career has focused on vertebrate evolution, the evolution of
birds, and the tempo and mode of the evolution of modern groups of
birds. His interest in the origin of birds blossomed in the late 1970s
when he wrote The Age of Birds for Harvard University Press.
In 1973 he wrote a rebuttal to the theory of hot-blooded dinosaurs in
the Journal of Evolution, and since that time has been
involved in the debate on bird origins. His latest book, The Origin
and Evolution of Birds (Yale University Press, 1999) treats many
of the main issues in the bird evolution controversy, and takes what he
calls the "ornithological" position, that is, that birds evolved flight
from the trees down, and from a common ancestor with dinosaurs, but not
directly from them.
Alan's interest in birds began as a teenager. Later, as an
undergraduate student at Louisiana State University, he had the good
fortune to participate in "bird" expeditions to Honduras, El Salvador,
and Peru. He has maintained an interest in Neotropical birds ever
since.
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John W. Fitzpatrick, Cornell University
CHAPTER 10: Bird Conservation
John W. Fitzpatrick (Ph.D., Princeton University, 1978) is
Director of the Cornell Laboratory of Ornithology, where he arrived in
September 1995. He was Executive Director of Archbold Biological
Station, a private ecological research foundation in central Florida,
from 1988 through Aug. 1995, and was Curator of Birds at the Field
Museum of Natural History (Chicago) from 1978 to 1989. He is a Fellow
of the American Ornithologists' Union (AOU). In 1985 the AOU awarded
him its highest research honor (Brewster Award), for his book and
numerous research articles (co-authored with Glen Woolfenden) on
demography, social behavior, and conservation of the endangered Florida
Scrub-Jay.
Fitzpatrick has led many expeditions to remote areas of South America,
especially the western Amazonian basin and the Andean foothills. He has
published numerous papers on Neotropical birds, including descriptions
of seven bird species new to science. Co-Author of the book,
Neotropical Birds: Ecology and Conservation (Univ. Chicago
Press, 1996), he has been engaged in applying science to real world
conservation issues throughout his career. Most recently, he helped
design and implement a major network of ecological preserves in central
Florida by convening panels of scientific experts and by engaging
county, state, and federal agencies, nongovernmental organizations, and
private industry in the process.
He serves on the national governing boards of The Nature Conservancy,
the National Audubon Society, and the Center for Biodiversity and
Conservation at the American Museum of Natural History. He is on two
Endangered Species Recovery Teams, including that of the world's rarest
bird, the Hawaiian Crow. He enjoys watercolor painting, has been a
birdwatcher since kindergarten.
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J. B. Heiser, Cornell
University
CHAPTER 4: What's Inside: Anatomy and Physiology
J. B. Heiser is a Senior Lecturer in the Department of
Ecology and Evolutionary Biology at Cornell University. A vertebrate
evolutionary ecologist, J. B. received his B.S. from Purdue Univerisity
and his Ph. D. from Cornell University. For the past 35 years has
taught a variety of courses in vertebrate comparative anatomy and
ecology.
J. B. began teaching on the Ithaca campus, but soon was teaching field
courses at the Shoals Marine Laboratory, an isolated island facility in
the Gulf of Maine that is cooperatively run by Cornell University and
the University of New Hampshire. Eventually he became director of the
Shoals program, a position that he held for 15 years. He won the Clark
Award for distinguished teaching and has traveled to every continent
and ocean to teach natural history "on location." Although J. B. was
trained as a marine biologist and his research focuses on the evolution
and interrelationships of coral reef fish, he has considerable natural
history experience in tropical forests worldwide. In his travels he has
watched birds (and fish) in every major biome and biogeographic region
that the planet has to offer, but he still gets a thrill out of
backyard birding in Upstate New York.
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Stephen W. Kress, Audubon
CHAPTER 2: A Guide to Bird Watching
Stephen W. Kress is Vice President for Bird Conservation for
the National Audubon Society, Manager of the Society's Maine Coast
Seabird Sanctuaries, and Director of the Seabird Restoration Program's
Project Puffin. He
also is ornithology program director for the Audubon Camp in Maine, an
adjunct faculty member in the Wildlife Department at the University of
Maine, Orono, and a Visiting Fellow at the Cornell Lab of Ornithology,
where each spring he teaches a popular course called Spring Field
Ornithology.
As director of Audubon's Seabird Restoration Program, Steve advises
and manages the development of techniques for re-establishing various
Maine seabird colonies, such as Atlantic Puffins, Leach's Storm-Petrel,
and Arctic, Common, and Roseate Terns. In the Pacific region, he has
studied the role of vocalizations in attracting endangered Dark-rumped
Petrels to artificial burrows in the Galápagos Islands, and
Short-tailed Albatross to decoys on Midway Island. He is author of many
books, including The National Audubon Society's Birder's Handbook,
The Bird Garden, and Project Puffin, as well as the Golden
Guide to Bird Life. He also has authored numerous scientific
papers on seabird biology and conservation.
During most of the year Steve lives on 33 acres of woods and meadows
near Ithaca, New York, where he manages his land for songbirds and
works on methods for restoring populations of Northern Bobwhite Quail.
He spends summers on the Maine coast, continuing his lifelong interest
in restoring nesting seabird colonies.
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Donald E. Kroodsma, University of
Massachusetts
CHAPTER 7: Vocal Behavior
Birds enter our lives in different ways. Some of us have been
bird-crazy as long as we can remember, but others discovered birds
later. I was a late-comer, as birds grabbed me from the chemistry lab
during my last year of college. How grateful I am that they have never
let go. Immediately after college I took two summer bird courses from
the famed Olin Sewall Pettingill, who was then director of the Cornell
Lab of Ornithology but taught at the University of Michigan field
station in Pellston, Michigan. He put a tape recorder, headphones, and
parabolic microphone in my hands and told me to "go out and tape record
some birds." In doing so, he changed my life. I went to graduate school
at Oregon State to study how young wrens learn their songs; and then
for eight years I had the good fortune to work with Peter Marler at
Rockefeller University in New York, on all aspects of bird song. The
time since college has been spent reveling in the who, what, when,
where, how, and why of bird song.
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Kevin J. McGowan, Cornell
University
CHAPTER 1: Introduction: The World of Birds
Kevin J. McGowan is the instructor of the Home Study Course
in Bird Biology and the new online course Courtship and Rivalry in
Birds at the Cornell Laboratory of Ornithology. He received his B.S.
and M.S. degrees in Zoology from the Ohio State University and a Ph.D.
in Biology from the University of South Florida, where he studied the
social development of Florida Scrub-Jays. He came to Cornell University
in 1988 as curator of the Ornithology and Mammalogy collections in the
Department of Ecology & Evolutionary Biology. In addition to caring
for the collections he conducted research on crows and taught classes
in specimen preparation, field collecting, the relationships of birds,
and Neotropical canopy biology. He moved to the Lab in 2001. He
helped create the original All About Birds web site and wrote the Bird
Guide section. He co-edited and wrote much of The Second Atlas of
Breeding Birds in New York State.
Kevin's primary research focuses on behavioral ecology of birds.
Currently he is studying the reproductive and social behavior of two
crow species in central New York state and investigating the impact of
West Nile virus on crow populations. He is an Elected Member of the
American Ornithologists' Union, director and webmaster for the
Federation of New York State Bird Clubs, and a member of the New York
State Avian Records Committee. He was former Secretary of the
Ornithological Societies of North America (OSNA), and former editor of
the Ornithological Newsletter, a bi-monthly OSNA
publication.
An avocational birder since childhood, Kevin has traveled throughout
North America and to Europe, Africa, and Central and South America to
watch and study birds.
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Daniel Otis, Cornell University
Birds and Humans: A Historical Perspective
Daniel Otis grew up on a farm in upstate New York. Since 1988
he has worked as a freelance editor, proofreader, and writer for Living
Bird and other Lab of Ornithology projects. Currently a Ph.D. student
in horticulture at Cornell, his research focuses on conservation of the
world's maple species and on assessing the extent and effects of Norway
maple invasiveness on northeastern forests.
Although plants rather than birds became his main interest, his
appreciation of nature derives partly from memorable experiences with
birds: "Once when I was a teenager, on a stormy November day, as I
walked along a hedgerow," he writes, "I was astonished to come across
an isolated little tree crowded with silent birds I'd never seen
before—Cedar Waxwings. Too exhausted to fly, they merely shuffled down
their perches a bit when I came close. And in March of some years, the
air was alive day and night with the sound of immense flocks of Canada
geese flying between the lake and the muddy cornfields around our
house." Experiences such as these, he notes, quicken one's appreciation
of the vivacity and fascination of the natural world, and help us
understand what conservationists are fighting for.
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Sandra G. Podulka,
Cornell University
Birds and Humans: A Historical Perspective
Sandy Podulka received a B.S. in Wildlife Biology from
Cornell University and an M.S. in Zoology (Animal Behavior) from the
University of Maryland, where she studied the function of song
repertoires in Song Sparrows. After graduate school, Sandy was a
Research Technician at Cornell for Dr. Stephen T. Emlen, analyzing the
social behavior of White-fronted Bee-eaters, but discovered she wanted
to spend more time sharing her love of nature with others, so turned
toward environmental education. She worked at the Cayuga Nature Center
for several years, and as an Adjunct Professor of Biology at Tompkins
Cortland Community College from 1988 to 1996, teaching courses in
biology and conservation. Since 1986, she has worked at the Cornell Lab
of Ornithology, in research, writing, public education, and
editingÛmost recently as one of the editors of the Home Study Course in
Bird Biology.
Sandy spent her childhood knee-deep in muddy ponds trying to catch
tadpoles and frogs, and roaming fields collecting butterflies—her first
real love. She has enjoyed birds as long as she can remember, but they
did not take center stage until she took a summer ethology course at
Cornell University from Dr. Bill Dilger. He hauled his students out
before dawn every morning and taught them to recognize birds by their
songs, and Sandy has been listening to them ever since. She has
participated in Christmas Bird Counts and Breeding Bird Atlases, and
has traveled to Costa Rica, Belize, Trinidad, and Peru to watch birds,
but her favorite bird watching site is her yard—which overlooks a
beaver pond with an ever-changing cast of avian actors.
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N. John
Schmitt
Illustrations
N. John Schmitt is a wildlife illustrator with a lifelong
interest in birds. Since leaving the United States Army in 1973 John
has devoted his life to a variety of ornithology related endeavors
which have taken him to many countries in Latin America, Asia, and
Europe. He has worked as a field biologist for both the Peregrine
Falcon and California Condor recovery programs.
John's work as a field biologist led to more serious devotion to
illustrating birds. He has illustrated several books, including the
National Geographic Society's 3rd edition of Birds of North
America; Clark's A Field Guide to the Raptors of Europe, the Middle
East, and North Africa; and Skutch's Birds Asleep. Currently, he
is working on bird field guides for Peru and India. John is a
self-taught taxidermist/museum preparator whose work is on display in
several California museums. He also co-leads ecotours in the United
States and abroad.
A native of California, John is an avid bird watcher. His time in the
field provides valuable inspiration and is integral to maintaining his
enthusiasm. He considers his notebooks, which he fills with written and
sketch notes, as a source of his most valuable reference and
inspiration. John encourages everyone interested in natural history to
keep a notebook.
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Stanley A. Temple, University of Wisconsin
CHAPTER 9: From Individuals to Ecosystems: Understanding Bird
Ecology
Stanley A. Temple is the Beers-Bascom Professor of
Conservation in the Department of Wildlife Ecology at the University of
Wisconsin in Madison. He is also the Chair of the graduate program in
Conservation Biology and Sustainable Development in the Institute for
Environmental Studies at Madison. A quintessential Cornellian, Stan
earned his B.S., M.S., and Ph.D. all at Cornell, and for years has
served as a member of the Lab of Ornithology's Administrative
Board.
Stan's professional activities focus on avian ecology and bird
conservation, with a special emphasis on endangered species. He and his
students have worked with some of the world's most endangered birds,
including Peregrine Falcons, California Condors, Whooping Cranes, and
dozens of endangered endemic species on islands around the world. To
date, none of those species has become extinct, and most are doing
significantly better as a result of his work.
Stan has been interested in birds as long as he can remember. Birds of
prey have always been among his favorites, and he has been a falconer
for 45 years. He feels fortunate to have incorporated all of his
ornithological pleasures into his professional life.
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David W. Winkler, Cornell University
CHAPTER 8: Nests, Eggs, and Young: The Breeding Biology of Birds
David Winkler was born and raised in Sacramento, California.
Unlike the rest of his family, he was a naturalist from about age four.
After a progression of enthusiastic interest in butterflies,
wildflowers, and herps, he finally settled on birds in his early teens.
He learned and studied local birds alone for two years, when, at his
first Christmas Count, he ran into Rich Stallcup. David then spent the
last two years of high school learning a great deal from Rich about the
birds of California. While in high school, an American Birds
article introduced David to The Herring Gull's World by Niko
Tinbergen, and a senior English project on gull taxonomy followed
within a couple of years. By his freshman year at U. C. Davis, David
was dreaming about all the species he might some day study as opposed
to seeing for his list. While attending U. C. Davis, David and friends
secured National Science Foundation funding for an ecological study of
Mono Lake. This cemented his attachment to the Mono Basin and its bird
life, and his friendship with David Gaines, with whom he co-founded the
Mono Lake Committee in 1978. A dissertation on the clutch sizes of
California Gulls (with Frank Pitelka at Berkeley) at Mono and Great
Salt Lakes was followed by post-docs at University of Gothenburg,
Sweden (with Malte Andersson); Oxford University (with John Krebs); and
Cornell University (with Paul Sherman). David joined the Cornell
faculty in 1988 upon the retirement of Tom Cade, and his research on
swallows world-wide, and the Ithaca Tree Swallows in particular, has
thrived ever since.
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